Floyd Gerald Werner 1921-1992.
Psyche 101(1-2):1-12, 1994.
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1.101 1994 NO. 1-2
Floyd Gerald Werner, 1 99 1
FLOYD GERALD WERNER 192 1-1992
^loyd Gerald Werner was born June 1, 192 1, to Frank and Edith mper Werner. He attended the public schools in Ottawa, Illinois here his career in insects began with his first publication in partment of Entomology, University of Arizona, Tucson, AZ 85721 uscrim received 7 December 1993,
2 Psyche [vo~. 101
1938, an account of the habitat and behavior of Doru aculeatum (Scudder), an earwig found in the marshes of northern Illinois. His love of science was fostered by a high school teacher, Charles J. Alikonis.
Upon graduation Floyd enrolled at Harvard College. In 1943, he was awarded a bachelor's degree in Biology, magna cum laude. At this time he was initiated into Phi Beta Kappa and Sigma Xi. Dur- ing these undergraduate years Floyd served as a student assistant in the Coleoptera Section of the Museum of Comparative Zoology, where he discovered many treasures that he would work on later in his career.
World War I1 interrupted Floyd's college career, giving him the opportunity to serve as an entomologist with the U.S. Army in the South Pacific. His medical survey unit worked with mosquitoes and malaria in Okinawa and Korea. In 195 1 he returned to Okinawa at the request of the Pacific Science Board to study sweet potato pests, as sweet potatoes were the chief source of carbohydrates for a pop- ulation still in post-war recovery.
In 1946-47, Floyd returned to the South Pacific as part of a sci- entific expedition charged with surveying the flora and fauna of the Philippine Islands, particularly Luzon, Mindanao and Palawa. The expedition was sponsored by the Field Museum of Natural History, Chicago and the Philippine Bureau of Science. Collections included everything from bizarre mayflies in the family Prosopistomatidae to giant fruit eating bats infested with Nycteribiidae to hoodless cobras. Stories about these times always brought a sparkle to Floyd's eyes.
Returning to Harvard in 1947 after these adventures, Floyd worked on a doctorate in Zoology. His specialty was insect taxon- omy, in particular the Meloidae and Anthicidae. Many papers on these families followed, climaxed by his work on the meloids of Arizona, published in 1966, and several major generic revisions of anthicids. Floyd and fellow student William L. Nutting took long collecting trips in the summers of 1948 and 1949 across the United States, including side trips into Canada and Mexico. They spent several months in the Southwest. A white bread truck, Floyd's infamous collecting vehicle, took him and Bill into many grand collecting spots, including one in South Dakota that turned out to be accessible only by a horse cart with old car tires, a surprise that left these compadres looking for a tow out. In 1950, Floyd was
19941 Olson 3
awarded his Ph. D. in Zoology. His dissertation was entitled "Stud- ies of Nearctic Anthicidae (Coleoptera)." Floyd's first job was as Assistant Professor of Zoology at the University of Vermont in 1950. It was here that he met his future wife, Frances Watson, who was also teaching in the Zoology Department. This union produced three children, Susan, William and John. All three are commissioned law officers working for wildlife agencies in Colorado or Arizona. They are carrying on their father's interest in and love of the great outdoors. In 1954, Floyd returned to the scene of those monumental trips of the late forties. He was hired by the University of Arizona ento- mology department, headed by Dr. Lawrence Carruth, to be part of the staff of the Arizona Economic Insect Survey, to expand the departmental insect collection as curator and to teach systematics and other courses as needed. Collecting emphasis was on insects important to the expanding agricultural community. He was pro- moted to Associate Professor in 1958 and Professor in 1962. At the University of Arizona, Floyd built the research collection from one Cambridge Unit that held 28 Schmitt boxes into the finest Southwest insect collection in the United States, with well over half a million specimens and more than thirteen thousand species of Arizona insects. The spectacular array of Coleoptera is the high point. Floyd, though, was the collector's collector, and his forays through Arizona with George Butler and Bill Nutting in the early fifties and sixties added many representatives of Hymenoptera and Diptera, including parasites valuable for biological control. His responsibilities in the department led to publications on scarab beetles and white grubs, parasites of cotton insects and other insects important to agriculture in Arizona. Studies made by his graduate students included bee-meloid interactions, range plant- insect associations, spiders as predators in crops and various taxo- nomic treatments.
His true love still lay with the small beetles in the families Anthicidae and Aderidae. In 1963 a sabbatical in South America with stops in Brazil and Argentina helped provide material for his comprehensive study of the genus Acanthinus in the New World. Other studies on anthicids included a revision of the genus Anthi- cus in North America and a preliminary study of the anthicids of New Zealand.
4 Psyche [vo~. 101
Floyd's final works included a revision of the Aderidae, a study made possible by the excellent collections of Karl Stephan, and a special paper on the genus Elonus dedicated to Dr. Frank M. Car- penter, Floyd's mentor while at Harvard. Floyd was also working on the aderids of New Zealand.
After his retirement in 1989, Floyd took on the unenviable task of Editor of the Coleopterist Bulletin, which he fulfilled to the end, his final volume of the Bulletin appearing the day after his death. He prided himself in making sure a job he took on was completed in a timely and professional manner.
Because being editor of a journal and continuing work on beetles wasn't enough, Floyd accepted the challenge of writing a book on the arthropods of the Southwest, for lay people to use to understand what Floyd enjoyed most, the world of insects. This book should be on the market in the spring of 1994.
Floyd was a member of many professional societies and served on a variety of their committees over the years. He was chairman of the Committee on Common Names of Insects and edited the 1982 revision for the Entomological Society of America. He was partic- ularly interested in the survival of taxonomy and the continuation of historical collections. He had visited a number of the collections of the world including those in London, Budapest and Paris, and contributed his curatorial skills and knowledge of beetles wherever possible at those institutions.
Some of the statistics on Floyd's professional accomplishments include 100 professional papers on a variety of insects, from bee- tles to mantispids, parasitic wasps to spiders. His knowledge of the arthropod world was truly awesome. He described 164 species of meloids, anthicids, aderids and cebrionids, and three new genera in the former three families. His name adorns such insects as an Ari- zona stonefly, numerous meloids, anthicids and many other taxa. As a citizen of Tucson, Floyd was active in groups such as the Tucson Men's Garden Club, Tucson Cactus and Succulent Society, the Tucson Natural History Society and many others. He was asked to speak before many local groups about insects. He was an origi- nal faculty member for the Tucson Audubon Society's Desert Ecol- ogy Institute.
Floyd loved to hunt and fish. He spent much of his leisure time outdoors experiencing the Arizona desert as a naturalist. He fished for sunfish and bass, bow-hunted for elk, antelope and javelina, and stalked the elusive black-tailed jackrabbit.
19941 Olson 5
Floyd's lasting obsession was his collection of Euphorbiaceae that occupied several greenhouses and a great part of the acre of yard around the Werner casa. Floyd started many rare plants from seed, priding himself in this conservation effort. He finally got to see euphorbias first hand in the wild when he traveled to South Africa with his wife and the Nuttings to attend the annual meeting of the Succulent Society in Johannesburg in 199 1. It seems quite amazing that Floyd waited until retirement to suddenly be afflicted with very untimely ills. He was hunting squirrels the weekend before he needed his gall bladder removed. This little incident caused him to miss a couple weeks of his last systematics class, a rare occurrence indeed. A year later he under- went triple-bypass heart surgery, another 'minor' setback for him. He came out of those like the stout-hearted German he was, and went back after life's challenges. His last battle, though, with can- cer was a losing one and Floyd died on December 20, 1992. It seems only appropriate to end this history of my mentor, col- league and good friend with an archygram from Floyd's favorite philosopher, Archy the Cockroach.
'insects have their own point
of view about civilization a man
thinks he amounts to a great deal
but to a flea or a mosquito
a human being is merely something
good to eat'
the lives and times of archy and mehitabel by don marquis
Werner, F. G. 1938. A report on the earwig Doru aculeatum (Scudder), from a marsh in Northern Illinois. Trans. Illinois State Acad. Sci. 31: 249. -- . 1943a. A revision of the genus Pleuropompha LeConte (Col., Meloidae). Psyche 50: 30-3.
-- . 1943b. Three new species of Cebrio (Col., Cebrionidae). Psyche 50: 34-6. -- . 1944. Some new North American species of Epicauta (Col., Meloidae). Psyche 50: 65-73.
-- . 1945. A revision of the genus Epicauta in America north of Mexico (Col., Meloidae). Bull. Mus. Comp. Zool. 95: 421-517.
6 Psyche [vo~. 101
-- . 1948. A note on the type specimen of Bagous sellatus LeConte (Col., Cur- culionidae). Psyche 54: 262.
Werner, F. G., and R. L. Edwards. 1948. Leptinus americanus LeConte taken on a shrew (Col., Leptinidae). Psyche 55: 5 1-4. Werner, F. G. 1949a. Epicauta diversicornis and its allies in the Neotropical Region (Col., Meloidae). Psyche 56: 74-80.
-- . 1949b. Additions to Epicauta, with new synonymy and a change of names (Col., Meloidae). Psyche 56: 93-1 11.
-- . 1951. Additions to the Nearctic Meloidae (Col.). Psyche 57: 131-6. --- . 1954a. Further notes on North American Epicauta, with new synonymy (Col., Meloidae). Psyche 60: 105-14.
--- . 1954b. A review of the subgenus Gnathospasta of the genus Epicauta (Col., Meloidae). Coleop. Bull. 8: 25-7. -- , 1954c. Pyrota plagiata (Haag) a valid Mexican species (Col., Meloidae). Bull. Brooklyn Entomol. Soc. 49: 102-4.
--- . 1955. Studies in the genus Epicauta of the North American Continent (Col., Meloidae). I. The Caviceps-Group. Bull. Brooklyn Entomol. Soc. 50: 1-12.
. 1956. Two new species of Emelinus from Arizona (Col., Aderidae). Psyche 63: 30-6.
Butler, G. D., and Werner, F. G. 1957. The syrphid flies associated with Arizona crops. Arizona Agric. Exp. Sta. Tech. Bull. 132, 12 pp. Werner, F. G. 1957a. Lappus thican$ormis, a new species from Michoacan. (Col., Anthicidae). Coleop. Bull. 10: 87-9.
--- . 1957b. A new species of Epicauta from the Gulf Coast of Texas (Col., Meloidae). Proc. Entomol. Soc. Washington. 59: 97-8. --- . 1957c. Two cases of intestinal myiasis in man produced by Hermetia (Diptera, Stratiomyiidae). Psyche 63: 112. Werner, F. G., and Butler, G. D. 1958. The reduviids and nabids associated with Arizona crops (Hemiptera). Arizona Agric. Exp. Sta. Tech. Bull. 1 33, 12 pp. Werner, F. G. 1958a. A revision of the Nearctic species of Tomoderus (Col., Anthi- cidae). Psyche 64: 51-9.
. 1958b. A new genus and species of Anthicidae from western United States (Col.). Psyche 64: 97-101.
-- . 1958c. Epicauta dugesi a valid species (Col., Meloidae). Psyche 64: 107-8. --- . 1958d. Some notes on Boheman's Anthicidae from "California" (Col.). Proc. Entomol. Soc. Washington 60: 2 13-1 6. --- . 1959. Studies in the genus Epicauta of the North American Continent (Col.). 11. The Uniforma-Group. Coleop. Bull. 12: 1-19. Butler, G. D.; Todd, F. E.; McGregor, S. E.; and Werner, F. G. 1960. Melissodes bees in Arizona cotton fields (Hymenop., Apidae). Arizona Agric. Exp. Sta. Tech. Bull. 139, 11 pp.
Werner, F. G. 1960a. A new character for the identification of the boll weevil and the thurberia weevil (Col., Curculionidae). Ann. Entomol. Soc. America 53: 548-9.
--- . 1960b. Hilarocassis exclamationis (L.), a tortoise beetle not previously reported from the United States (Col., Chrysomelidae). Coleop. Bull. 14: 94. . 1960c. Carpophilus longiventris in saguaro blossoms (Col., Nitidulidae). Psyche 66: 35-6.
. 1960d. Emelinus melsheimeri (Lee.) in Arizona (Col., Aderidae). Psyche 66: 36.
. 196la. A note on the prey and a nesting site of Cerceris truncata Cameron (Hymenop., Sphecidae). Psyche 67: 43-4.
--- . 1961b. A revision of the genus Vacusus Casey (Col., Anthicidae). Ann. Entomol. Soc. America 54: 798-809.
. 1961c. Anthicus tobias Marseul, another tramp species (Col., Anthicidae). Psyche 68: 70-2.
Butler, G. D., and Werner, F. G. 196la. Pentatomids associated with Arizona crops (Hemip.). Arizona Agric. Exp. Sta. Tech. Bull. 140, 16 pp. --- . 1961b. The distribution and host plants of May beetles in Arizona (Col., Scarabaeidae). Arizona Agric. Exp. Sta. Tech. Bull. 147, 19 pp. Werner, F. G. 1962a. The first instar larva of Epicauta, subgenus Gnathospasta (Col., Meloidae), Verhandl. XI. Int. Kongress Entomol. Wien 1960, vol. 1: 106-9.
. 1962b. A redefinition of Acanthinus as an important element in the anthicid fauna of the Neotropical Region (Col., Anthicidae). Verhandl. XI. Int. Kongress Entomol. Wien, vol. 1 : 109-12.
-- . 1962c. The Aderidae of Bata Caves, Malaya (Col.). Pacific Insects 4: 121-7. --- . 1962d. The species of Mecynotarsus related to elegans LeConte (Col., Anthicidae). Proc. Entomol. Soc. Washington. 64: 79-86. -- . 1962e. A revision of the Nearctic species of Sapintus (Col., Anthicidae). Ann. Entomol. Soc. America 55: 492-8.
-- . 1964. A revision of the North American species of Anthirus, s. str. (Col., Anthicidae). Misc. Publ. Entomol. Soc. America 4: 139-242. Werner, F. G., and Butler G. D. 1965. Some notes on the life history of Plega banksi (Neurop., Mantispidae). Ann. Entomol. Soc. America 58: 65-8. Butler, G. D., and Werner, F. G. 1965. Light-trap records of three cactus-eating moths in Arizona (Lepidop., Pyralidae). Pan-Pac. Entomol. 41: 10-12 Werner, F. G. 1965a. A key to the described species of New World Anthicidae of the genus Acanthinus LaFert6, with new synonymy (Col.). Pap. Avuls. Dept. Zool., Secr. Agric. Sao Paulo 18 (Article I): 1-15. --- . 1965b. Insects of Micronesia, Coleoptera: Anthicidae. Insects of Microne- sia 16: 255-69.
-- . 1965c. Family Anthicidae, pp. 122-9. In Hatch, M. H. The Beetles of the Pacific Northwest, Part IV. U. Washington Pubs. Biol. 16. U. Washington Press, Seattle.
--- . 1966a. Notes on the South American species of Vacusus, with a new species and two new synonyms (Col., Anthicidae). Ann. Entomol. Soc. America 59: 218-22.
-- . 1966b. A new genus of Anthicidae from Chile (Col.), Ann. Entomol. Soc. America 59: 222-6.
-- . 1966c. A redefinition of Ischyropalpus, and six new species (Col., Anthici- dae). Psyche 72: 191-209.
-- . 1966d. Anthicus postsignatus and similar species in southern South Amer- ica (Col., Anthicidae). Ann. Entomol. Soc. America 69: 362-8. -- . 1966e. A preliminary account of the Anthicidae of Venezuela (Col.). Rev. Fac. Agron., Univ. Central Venezuela 3(4): 9-23. -- . 1966f. A revision of Acanthinus (Col., Anthicidae). I. The Bimaculifer- Group. Ann. Entomol. Soc. America 59: 509-13.
8 Psyche [vo~. 101
-- . 1966g. A revision of Acanthinus (Col., Anthicidae). 11. The Angusticollis- Group. Ann. Entomol. Soc. America 59: 746-5 1. -- . 1966h. A revision of Acanthinus (Col., Anthicidae). 111. The Spinicollis- Group. Ann. Entomol. Soc. America 59: 1267-76. Butler, G. D.; Werner, F. G.; and Levin, M. D. 1966. Native bees associated with safflower in south central Arizona. (Hymenop., Apoidea). Jour. Kansas Entomol. SOC. 39: 43-6.
Werner, F. G.; Enns, W. R.; and Parker, F. H. 1966. The Meloidae of Arizona (Col.). Arizona Agric. Exp. Sta. Tech. Bull. 175, 96 pp. Werner, F. G. 1967a. A revision of Acanthinus (Col., Anthicidae). IV. Ann. Ento- mol. Soc. America 60: 225-73.
--- . 1967b. The zoological results of Gy. Topal's collectings in South Argentina. 21. Coleoptera: Anthicidae. Acta Zool. Acad. Scientiarum Hungari- cae 13: 237-43.
--- . 1967c. A key to the Anthicidae of Hawaii, with one new species (Coleoptera). Proc. Hawaiian Entomol. Soc. 19: 310-16. --- , 1967d. A revision of Acanthinus (Coleoptera : Anthicidae). V. The Stri- atopunctatus Group and some related forms. Ann. Entomol. Soc. America 60: 535-49.
--- . 1967e. A revision of Acanthinus (Col., Anthicidae). VI. Ann. Entomol. Soc. America. 60: 1217-34.
Butler, G. D.; Ritchie, P. L.; and Werner, F. G. 1968. The effect of temperature on the life cycle of the alfalfa seed chalcid and its parasites. Arizona Agric. Exp. Sta. Tech. Bull. 185- 17 pp.
Werner, F. G. 1968. The Arizona brown spider. Progr. Agric. Arizona 20: 12-13. -- . 1969. Two flights of Scaptolenus (Col., Cebrionidae). Coleop. Bull. 23: 26-7. Humphrey, R. R., and Werner, F. G. 1969. Some records of bee visitations to the flowers of Idria colurnnaris. J. Arizona Acad. Sci. 5: 243-4. Werner, F. G. 1970a. A revision of Acanthinus (Col., Anthicidae). VII. Ann. Ento- mol. Soc. America 63: 111-28.
-- . 1970b. A revision of Acanthinus (Col., Anthicidae). VIII. The Australian species. Ann. Entomol. Soc. America 63: 486-90. --- . 1970c. A revision of Acanthinus (Col., Anthicidae). IX. The Leporinus, Scitulus, Cuyabanus and Myrmecops-Groups. Ann. Entomol. Soc. America 63: 718-31.
-- . 1970d. A revision of Acanthinus (Col., Anthicidae). X. Albicinctus-Group and conclusion. Ann. Entomol. Soc. America 63: 899-76. --A . 1970e. Terrestrial insects of the Rita Blanca Lake deposits, pp. 123-130, pi. 16-20. In: Anderson, R. Y. and D. W. Kirkland. Paleoecology of an early Pleistocene lake on the High Plains of Texas. Mem. Geol. Soc. America 113, 215 pp.
Rosander, R. W., and Werner, F. G. 1970. Larvae of some Arizona Species of Phyl- lophaga (Col., Scarabaeidae). Ann. Entomol. Soc. America 63: 1136-42. Drake, J. L.; Ware, G. W.; and Werner, F. G. 1971. Insecticidal effects on soil arthropods. J. Econ. Entomol. 64: 842-5. Werner, F. G. 1973. Revision of the Nearctic Ischyropalpus (Col., Anthicidae). Ann. Entomol. Soc. America 66: 1055-64.
--- . 1974a. Three new Epicauta from Mexico (Col. Meloidae). Proc. Entomol. Soc. Washington 75: 458-63.
19941 Olson 9
. 1974b. Further description of Epicauta prosopidis Werner (Col., Meloidae). Proc. Entomol. Soc. Washington 76: 40.
-- . 1974c. A new genus of primitive Meloidae from West Texas. (Col.). Psy- che 8 1 : 147-54.
Erickson, E. H., and Werner, F. G. 1974a. Bionomics of Nearctic bee-associated Meloidae (Col.); life histories and nutrition of certain Meloinae. Ann. Entomol. Soc. America 67: 394-400.
-- . 1974b. Bionomics of Nearctic bee-associated Meloidae (Col.); life histories and nutrition of certain Nemognathinae. Ann. Entomol. Soc. America 67: 401-6.
--- . 1974c. Bionomics of Nearctic bee-associated Meloidae (Col.). A compara- tive analysis of larval host-seeking behavior among the Meloinae and Nemo- gnathinae. Ann. Entomol. Soc. America 67: 903-8. Werner, F. G. 1975a. A review of the Chilean Anthicidae (Col.). Rev. Chilena de Entomol. 8: 27-34.
--- . 1975b. New synonymy in the Nearctic Anthicidae. Proc. Entomol. Soc. Washington 77: 290.
--- . 1975c. Additions to the Nearctic Anthicus (Col., Anthicidae). Proc. Ento- mol. Soc. Washington 77: 472-7.
Erickson, E. H.; Enns, W. R.; and Werner, F. G. 1976. Bionomics of the bee-associ- ated Meloidae (Col.); bee and plant hosts of some Nearctic meloid beetles-a synopsis. Ann. Entomol. Soc. America 69: 959-70. Sluss, T. P.; Rockwood Sluss, E. S.; and Werner, F. G. 1977. Enzyme variation in semi-isolated populations of the mountain fly, Chamaemyia herbarum. Evolu- tion 3 1 : 302-1 2.
Mason, C. E., and Werner, F. G. 1978. Monitoring insect parasites in a cotton pest management program. Arizona Agric. Exp. Sta. Res. Report 276, 28 pp. Werner, F. G. 1978. Keys for the identification of parasitic insects in Arizona agri- cultural areas. Arizona Agric. Exp. Sta. Tech. Bull. 236, 38 pp. Werner, F. G.; Moore, L.; and Watson, T. F. 1979. Arizona Cotton Insects. Arizona Coop. Ext. Serv. Bull. A23R, 38 pp.
Hetz, M. W., and Werner, F. G. 1979a. Xeranobium laticeps (Col., Anobiidae) reared from Haplopappus tenuisectus (Compositae), with description of the larva. Proc. Entomol. Soc. Washington 81: 583-7. --- . 1979b. Insects associated with roots of some rangeland Compositae in southern Arizona. Southwestern Entomol. 4: 285-8. Werner, F. G. 1979c. Tachinid flies collected in a Phoenix, Arizona cotton field. Southwestern Entomol. 4: 282-4.
Hetz, M. W., and Werner, F. G. 1980. Descriptions of the larvae of two olethreutine moths reared from roots of woody Compositae. Ann. Entomol. Soc. America 73: 536-40.
Thomas, D. B., and Werner, F. G. 1981. Grass feeding insects of the western ranges: an annotated checklist. Arizona Agric. Exp. Sta. Tech. Bull. 243, 50 pp. Werner, F. G., editor. 1982. Common names of insects and related organisms 1982. Entomol. Soc. America, 132 pp.
Butler, G. D.; Henneberry, T. J.; Werner, F. G.; and Gillespie, J. M. 1982. Seasonal distribution, hosts, and identification of parasites of cotton insects. USDA, ARS, Agric. Rev. & Manuals: ARM-W-27, 54 pp.
Werner, F. G. 1983a. Neotropical Sapintus, with a general key to species (Col., Anthicidae). Proc. Entomol. Soc. Washington 85: 405-25.
10 Psyche [vo~. 101
--- . l983b. Anthicidae of the Greater Antilles, and a new species from Venezuela (Col.). Psyche 90: 21 1-25.
-- . 1986. Frank Henry Parker, 1910-1984. Pan-Pac. Entomol. 62: 1-5. Stanghellini, M. E.; Werner, F. G.; Turner, B. C.; and Watson, M. C. 1988. Seedling death of Salicornia attributed to Metachroma larvae. The Southwestern Ento- mologist. 13: 305.
Werner, F. C. 1990. Revision of the Aderidae of Eastern North America. J. New York Entomol. SOC. 98: 187-232.
-- . 1992. The Nearctic species of Elonus (Coleoptera: Aderidae) Psyche. 99: 245-264.
New genera and species of insects named by Floyd G. Werner Anthicidae
1) Acanthinus acutus
2) A. ambiguus
3) A. australiensis
4) A. bechyneorum
5) A. blackburni
6) A. bokermanni
7) A. bordoni
8) A. browni
9) A. ceibensis
10) A. continuus
11) A. cristatus
12) A. darlingtoni
13) A. diffusus
14) A. egleri
15) A. elegantulus
16) A. fairchildi
17) A. fastigatus
18) A. fimbriatus
19) A. formiciformis
20) A. freyorum
21) A. fucosus
22) A. geijskesi
23) A. glareosus
24) A. hapacarensis
25) A. harringtoni
26) A. imitans
27) A. invitus
28) A. klagesi
29) A. kraussi
30) A. lanceatus
31) A. lucidus
32) A. nevermanni
33) A. paraguayensis
34) A. parianae
35) A. pilositibia
36) A. pullus
37) A. rohweri
38) A. rosalesi
39) A. schwarzi
40) A. silvai
41) A. simplicisternum
42) A. solus
43) A. spinosior
44) A. veracruzensis 87) S. ovalis
45) A. zeteki 88) S. sentis
46) Anthicus antilleorum 89) S. similis
47) A. antiochensis
90) S. spinulosus
48) A. barbatus 91) S. subulatus
49) A. blackwelder 92) Tomoderus inhabilis 50) A. comanche 93) Vacusus martinsi
5 1) A. custodiae 94) V. vulgaris
52) A. darlingtoni
53) A. hispaniolae
54) A. hondurensis
55) A. margaritae
56) A. musculus
57) A. panamensis
58) A. potosianus
59) A. soledad
60) A. sonorensis
61) A. torquatus
62) Chileanthicus penai*
63) Ischyropalpus alvarengai
64) I. cochisei
65) I. dispar
66) I. decoratus
67) I. eryngii
68) I. freyi
69) I. gemellus
70) I. placidus
71) 1. puteifer
72) I. tibialis
73) Lappus thicaniformis
74) Mecynotarsus hispaniolae
75) Sapintus alvarengai
76) S. arizonicus
77) S. balteatus
78) S. canaliculatus
79) S. capitatus
80) S. caudatus
8 1) S. creber
82) S. curvipilosus
83) S. golbachi
84) S. lemniscatus
85) S. lobatus
86) S. malkini
1) Epicauta afoveata
2) E. alpina
3) E. andersoni
4) E. arizonica
5) E. aspersa
6) E. balli
7) E. barberi
8) E. bipunctata
9) E. bispinosa
10) E. brunnea
I 1) E. calcarata
12) E. californica
13) E. cicatrix
14) E. ennsi
15) E. ensiformis
16) E. ficta
17) E. floridensis
18) E. fortis
19) E. hubbelli
20) E. isthmica
21) E. kansana
22) E. laevicornis
23) E. lauta rossi
24) E. liebecki
25) E. nigritibialis
26) E. nogales
27) E. normalis
28) E. occidentalis
29) E. parkeri
30) E. phoenix
31) E. polingi
32) E. prosopidis
33) E. punctipennis
34) E. senilis
35) E. selanderorum
36) E. solani
37) E. tenebrosa
38) E. tenuemarginata
39) E. texana
40) E. triquetra
41) E. uniforma
42) E. ventralis
43) Lytta mirifica
44) L. navajo
45) Pleuropompha tricostata
46) Thambospasta howdeni*
1) Cebrio abnormis
2) C. atokanus
3) C. bruesi
1) Aderus mcclurei
2) Elonus chisosensis
3) E. excavatus
4) E. hesperus
5) E. simplex
6) Emelinus butleri
7) E. huachucanus
8) Euglenes batuensis
9) E. cephalicus
10) E. malayanus
11) Gymnoganascus stephani*
12) Vanonus balteatus
13) V. macrops
14) V. musculus
15) V. oklahomensis
16) V. uniformis
17) V. valgus
18) Zonantes floridanus
19) Z. mississippiensis
20) Z. ouachitanus
21) Z. pallidus
'"Indicates genus named by F. G. Werner
Volume 101 table of contents