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Ingolf S. Askevold.
An annotated list of Nearctic Donaciinae (Coleoptera: Chrysomelidae): the generic classification and type specimens of the New World species.
Psyche 98:165-192, 1991.

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AN ANNOTATED LIST OF NEARCTIC DONACIINAE (COLEOPTERA: CHRYSOMELIDAE): THE GENERIC CLASSIFICATION AND TYPE SPECIMENS OF THE NEW WORLD SPECIES.
Department of Entomology, University of Manitoba, Winnipeg, Manitoba. Canada R3T 2~2'
During systematic revision of Nearctic donaciines, the majority of type specimens were examined. Some lectotype and neotype designations, made where necessary in the context of revisions of genera (Plateumaris Thomson and Neohaemunia Szkkessy) or solution of specific taxonomic problems, were detailed elsewhere (Askevold 1987a,b, 1988, 1990, 1991). Some taxonomic problems remain despite relatively recent treatment by Marx (1957) of Donacia F. (sensu lato). I indicated earlier that Donacia is repre- sented in North America by 10 species in Donacia (s. str.), and 21 described species in Dunacia (Donaciomima) Medvedev, but details were not given then (Askevold 1990). The purpose of the present paper is to detail remaining lectotype designations among Neasctic members of the genus Donacia F., to list the Nearctic members of Donaciinae, and to detail relevant tax- onomic references and information (i.e., synonymies, taxonomic changes since prior authors, and data about type specimens). Schaeffer (1919, 1925) indicated only if he based his species on a type and allotype, but not that there were also many pasatypes; paratype information is included insofar as such specimens have been found. Species are listed within genera and subgeneia alpha- betically rather than phylogenetically, because phylogenies have been reconstructed only for members of Plateumaris and Neo- haemonia (Askevold 199 1, 1988).
Present address: Entomology-Biological Control, Division of Agricultural Sci- ences, Florida A & M University, Tallahassee, FL 32307-2001 Manuscript received June 7, 1991.




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166 Psyche [vo~. 98
The following list includes all described extant New World Donaciinae, serving to summarize the many taxonomic changes that have resulted from revision of North American Donaciinae. Details of historical use of species names is given where substan- tial changes in name use occurred [i.e., changes since Schaeffer (1925), Marx (1957) and Jolivet (1970)l. The taxonomic history of some names is complicated, often crossing generic boundaries; some species were synonymized with more than one taxon at once, based on composition of the original type series. I have judged that what is important is current synonymy and whether or not that dif- fers from the most recent uses of names, and therefore have not pried deeply into tracing the varied uses of names since their origi- nal description.
Data of individual labels borne by type specimens, where recorded, are stated verbatim and are separated by a slash ("I"). The museums and private collections in which type specimens of any kind are known to be preserved are identified by correspond- ing codens:
AMNH. Department of Entomology, American Museum of Natural History, Central Park West at 79th St., New York, NY 10024, U.S.A.
APHIS. Animal and Plant Health Inspection Service, USDA, Room 308, 40 South Gay St., Baltimore, MD 21202, U.S.A. BMNH. Department of Entomology, British Museum of Natural History, Cromwell Road, London SW7 5BD, England. CARR. J.L. and A.F. Carr, 24 Dalrymple Green, N.W. Calgary, Alta. Canada, T3A 1Y2.
CAS. Department of Entomology, California Academy of Sci- ences, Golden Gate Park, San Francisco, CA 941 18, U.S.A. CDAS. California Department of Food and Agriculture, California State collection of Arthropods, Insect Taxonomy Laboratory, 1220 N Street, Sacramento, CA 95814, U.S.A. CLCH. C. Chantal, B.P. 2072, St. Nicolas-est, P.Q., Canada, GOS 3L0.
CMP. Section of Entomology, Carnegie Museum of Natural His- tory, 4400 Forbes Ave., Pittsburgh, PA 15213, U.S.A.



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199 11 Askevold 167
CNC. Biosyntematics Research Centre, Agriculture Canada, Ottawa, Ont., Canada KIA OC6.
CUIC. Department of Entomology, Comstock Hall, Cornell Uni- versity, Ithaca, NY 14853, U.S.A.
DEFW. Department of Entomology, Fisheries and Wildlife, Uni- versity of Minnesota, 219 Hodson Hall, 1980 Folwell Ave., St. Paul, MN 55108, U.S.A.
EGRC. E.G. Riley, Department of Entomology, Texas A & M Uni- versity, College Station, TX 77843-2475, U.S.A. FMNH. Field Museum of Natural History, Roosevelt Road at Lake Shore Drive, Chicago, IL 60605, U.S.A.
FSCA. Florida State Collection of Arthropods, Division of Plant Industry, Florida Department of Agriculture, 19 1 1 S.W. 34th St., P.O. Box 1269, Gainesville, FL 32602, U.S.A. INHS. Department of Entomology, Illinois Natural History Survey, Natural Resources Bldg., Urbana, IL 6 180 1, U.S.A. ISAC. I.S. Askevold, Entomology-Biological Control, Division of Agricultural Sciences, Florida A & M University, Tallahas- see, FL 32307, U.S.A.
JBWM. J.B. Wallis Museum, Department of Entomology, Univer- sity of Manitoba, Winnipeg, Man. R3T 2N2, Canada. LEM. Collection Provancher, Dkpartement de biologie, Universite Laval, Quebec, P.Q., G1K 7P4, Canada.
LSU. Department of Entomology, Louisiana State University, Baton Rouge, LA 70803, U.S.A.
MCZ. Department of Entomology, Museum of Comparative Zool- ogy, Harvard University, Cambridge, MA 02138, U.S.A. MLU. Section Biowissenschaften, WB Zoologie, Martin-Luther- Universitat Halle-Wittenberg, Domplatz 4, 4010 Halle (Saale) Germany.
NMDC. N.M. Downie, 505 Lingle Terrace, Lafayette, IN 47901, U.S.A.
OSUC. Systematic Entomology Laboratory, Department of Ento- mology, Oregon State University, Corvallis, OR 9733 1, U.S.A.
PUR. Department of Entomology, Entomology Hall, Purdue Uni- versity, West Lafayette, IN 47907, U.S.A. ROM. Department of Entomology, Royal Ontario Museum, 100 Queen's Park, Toronto, Ont. M5S 2C6, Canada.



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168 Psyche [vo~. 98
SCSU. Department of Biological Sciences, St. Cloud State Univer- sity, St. Cloud, MN 56301, U.S.A.
SMCL. S.M. Clark, Pest Identification Laboratory, West Virginia Department of Agriculture, Charleston, WV 25305, U.S.A. TAMU. Department of Entomology, Texas A & M University, College Station, TX 77843-25 16, U.S.A.
UAE. Department of Entomology, University of Alberta, Edmon- ton, Alta. T6G 2E3, Canada.
UMHF. Universitetets Zoologiska Museum, Entomologiska Avdel- ningen, N. Jarnvagsgatan 13, SF-00100 Helsingfors 10, Fin- 1 and.
UMMA. Museum of Zoology, Insect Division, The University of Michigan, Ann Arbor, MI 48109, U.S.A.
USNM. National Museum of Natural History, Smithsonian Institu- tion NHB- 105, Washington, D.C. 20560, U.S .A. UVDZ. Department of Zoology, Marsh Life Science Bldg., Uni- versity of Vermont, Burlington, VT 05401, U.S.A. ZMHB. Zoologisches Museum, Museum fur Naturkunde der Hum- bolt-Universitat zu Berlin, Invalidenstr. 43, 1040 Berlin, Ger- many.
Plateumaris Thomson (1859:154). Revision of genus by Askevold (1 99 1). Rather than making numerous, repetitive statements about taxonomic details for this genus, users are referred to Askevold (1991) for details about type specimens. Unless other- wise stated, lectotype and neotype designations were made by Askevold (1991); a detailed account of the historical use of names is given there also, and not repeated here. P. aurifera (LeConte, 185 l:3 13). Lectotype (3 MCZ #4247. Vali- dated, and removed from synonymy with P. germari by Askevold (1991).
P. balli Askevold (1991). Holotype (7, CNC #19497, and 23 paratypes, ISAC, USNM, AMNH, CNC, APHIS, UAE, UVDZ, CAS and SMCL.
P. diversa (Schaeffer, 1925:143). Holotype Q, USNM #42397 and allotype (7, USNM.
P. dubia (Schaeffer, 1925: 152). Holotype (7, USNM #4240-C, allo- type Q USNM #42400 and 4 paratypes, USNM.



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199 11 Askevold 169
Donacia idola Hatch (1938: 110). Holotype 0, USNM type #103893, allotype Q OSUC and 45 paratypes, OSUC. Syn onymy with P. dubia by Askevold (1991).
P. flavipes (Kirby, 1837:223). Holotype Q, BMNH. This name was used incorrectly by previous authors; the taxon previously known as P. flavipes is now called P. shoemakeri (Schaef- fer). The taxon formerly called P. wallisi is now correctly called P. flavipes.
Donacia wallisi (Schaeffer, 1925: 147). Holotype 0 USNM #42399, allotype Q USNM #42399, and 13 paratypes, USNM. Synonymy with P. flavipes by Askevold (1 99 1). P. frosti (Schaeffer, 1925). Plateumaris frosti was formerly placed as a subspecies of P. emarginata (now P. nitida). Donacia emarginata var. frosti Schaeffer (1925:136). Holotype 0, USNM #42395, allotype Q, USNM #42395, 28 paratypes. USNM, and 3 paratypes, MCZ.
P. fulvipes (Lacordaire, 1845: 192). Holotype 0, BMNH. P. germari (Mannerheim, l843:306). Lectotype Q, MCZ #4250, and 1 syntype, UMHF.
Donacia flavipennis Mannerheim (1 843:306). Lectotype J, MCZ #425 1, and 1 syntype, UMHF. Specimens from western Canada possessing flavous elytra and/or pronoturn were given varietal status by Schaeffer (1925); Askevold (1991) gave no taxonomic status to specimens showing such varia- tion. Synonymy with P. germari by Askevold (1 99 1). Donacia dives LeConte (1 85 1 :3 14). Holotype Q, MCZ #4245. Donacia serricauda Schaeffer (l919:3 18). Holotype 9, USNM #42396.
P. metallica (Ahrens, 18 lO:33). Neotype 0, CNC #19493. Donacia femoralis Kirby (1837:223). Lectotype 9, BMNH. Donacia cataractae Newman (1 838:39 1). Lectotype 0, BMNH. Donacia parva Lacordaire (1 845 : 190), Lectotype 0, BMNH. Donacia indica Melsheimer (1 847: 158) (not Clark, 1 866: 1 ). Lectotype Q, MCZ #33363.
Donacia nana Melsheimer (1847:160). Neotype 0, MCZ #32953.
Donacia gentilis LeConte
(1 85 1 :3 14). Lectotype 0, MCZ
#4249, and two paralectotypes MCZ #4249.



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170 Psyche [vo~. 98
P. neomexicana (Schaeffer, 1925: 154). Holotype 0, USNM #424O 1 - 1, allotype Q USNM #424O 1, three paratypes, USNM, and two paratypes, MCZ #27227 [Fall colln.]. Donacia longicollis Schaeffer (1925:156). Holotype 0, USNM #4248-2, allotype USNM #42402 and 22 paratypes, USNM. Synonymized with P. neomexicana by Askevold (1991). Donacia vermiculata Schaeffer (1925: 158). Holotype 9, USNM. Synonymized with P. neomexicana given by Askevold (1991).
P. nitida (Germar, 181 1:31). Lectotype Q MLU. This name was used incorrectly by Schaeffer (1925); the species misidenti- fied by Schaeffer as P. nitida is now called P. schaefferi. The taxon to which the name P. nitida applies was called P. emarginata by Schaeffer (1925) and other authors. Donacia emarginata Kirby (1 837:224). Lectotype 0, BMNH. Synonymized with P. nitida by Askevold (1991). Donacia binodosa LeConte (1 85 1 :3 16) (cf. Suffrian, 1872). Unjustified emendation or nomen nudum. Synonymy with P. nitida by Askevold (1 99 1).
Donacia juncina Couper (1 864:88). Lectotype Q, LEM, desig- nated by Becker (1974). Synonymized with P. nitida by Askevold (1991).
Donacia emarginata var. pacifica Schaeffer (1 925: 135). Holo- type 0, USNM #42394, allotype Q, USNM #42394, and 9 paratypes, USNM. Synonymized with P. nitida by Askevold (1991).
P. notmani (Schaeffer, 1925: 132). Holotype 0, USNM #42393, allotype Q #42393, and five paratypes, USNM, 1 paratype MCZ #27226 [Fall Colln.].
P. pusilla (Say, 1827:293). Neotype (3, MCZ type #32950. Donacia rugifrons Newman (1838:391). Types of this name have not been found, but should be in BMNH. Three speci- mens (of P. rufa, however, not P. pusilla) were found in the MCZ that are labelled Trenton Falls, New York, collected by Doubleday. Similar labels are borne by specimens of two other Newman species - D. cincticornis and D. cataractae - and I consider the possibility that these might be specimens of Newman's D. rugifrons.




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19911 Askevold 17 1
Donacia pyritosa LeConte 1857(1860):66. Lectotype (7, MCZ type #4246. This name was formerly used as a variety of P. pusilla, but synonymized by Askevold (1991). P. robusta (Schaeffer, l919:3 18). This name was used as a variety of P. pusilla until elevated to species status by Askevold (1991).
Donacia pusilla var. robusta Schaeffer (1919:318). Holotype (7, USNM #42398, and 3 paratypes USNM #42398. P. rufa (Say, 1827:283). Neotype (7, MCZ #32947. This species was formerly called P. sulcicollis Lac. The name D. rufa has been used incorrectly by authors (Marx 1957:250, Schaeffer 1925: 117), and was transferred from Donacia to Plateumaris by Askevold (1991) [cf. also D. (Donaciomima) tuberculata Lacord., below].
Donacia affinis Kirby (1 837:224) (not Kunze, 18 18). Lectotype 9, BMNH. Transferred to synonymy with P. rufa from syn- onymy with P. sulcicollis by Askevold (1991). Donacia sulcicollis Lacordaire (1845:186). Lectotype 0, BMNH. Synonymized with P. rufa by Askevold (1991). Donacia chalcea Lacordaire (1 845: 188). Holotype 9, MLU. Transferred to synonymy with P. rufa from P. flavipes, by Askevold (1991).
Donacia kirbyi Lacordaire (1845:202). Replacement name for D. affinis Kirby; transferred to synonymy with P. rufa from P. sulcicollis by Askevold (1 99 1).
Donacia jucunda LeConte (1 85 1 :3 15). Holotype 0, MCZ #4248. Transferred to synonymy of P. rufa from P. sulcicol- Us by Askevold (1991).
P. schaefferi Askevold (1991). Holotype (7, USNM, and 7 paratypes, USNM, CUIC and UMMA.
Donacia nitida, sensu Schaeffer (1 925: 130) (see P. nitida, above).
P. shoemakeri (Schaeffer, 1925: 129). This species was formerly incorrectly known as P. flavipes (see P. flavipes, above), val- idated by Askevold (1 99 1).
Donacia flavipes var. shoemakeri Schaeffer (1925:129). Holo- type (7, USNM #4239-1 and 5 paratypes USNM #42391. Donacia flavipes var. lodingi Schaeffer (1 925: 129). Holotype 0, USNM #4239-2 and 1 paratype, USNM #42392. Southern



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172 Psyche [Vol. 98
specimens, from Alabama, were given varietal status by Schaeffer (1925), but Askevold (1991) did not give this geo- graphic variation taxonomic status.
Poecilocera Schaeffer (1919:316). Elevated to genus status by Askevold (1990)
P. harrisii (LeConte, 1851 :316). Lectotype Q MCZ #4244, desig- nated by Askevold (1990535); the abdomen is now missing but was present when I first examined it. LeConte stated he had at least two specimens, o and Q, although only a single specimen remains in the LeConte collection. P. harrisii was placed in Sominella by Goecke (1931), but Marx (1957) reverted to place- ment in Donacia (Poecilocera), evidently unaware of Goecke's treatment.
Domiella Reitter (1920:38). Elevated to genus by Askevold (1990), this name has been treated by European authors as a subgenus of Donacia, which has not been recognized in North America despite direct comparisons by Schaeffer (1925) with species occurring in the Palaearctic Region. D. pubicollis (Suffrian, 1872:21). Transferred to Donaciella by Askevold (1990). Suffrian did not indicate the number of specimens before him, but given a range of size measure- ments, had at least two. A series of 12 specimens are pre- served in MLU, associated with a Suffrian label. As determined previously (see P. chalcea and P. nitida, above) these specimens are also likely original Suffrian material, and are here designated. LECTOTYPE 0, following a Suf- frian label ["pubicollis m. Illinois", on large, green, black- bordered paper]: "25519 [red handwriting on small white label] 1 LECTOTYPE 0 Donacia pubicollis Suffrian 1872 design. I.S. Askevold 1991". The remaining 11 specimens are here designated paralectotypes. These bear the same red, handwritten number and my paralectotype label: 255 16 (cT), 25517 (o), 25520 (o), 25521 (o), 25522 (o), 25523 (Q), 25524 (Q), 25525 (Q), 25526 (Q), 33256 (Q) and 25518 (sex undetermined, abdomen missing).




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19911 Askevold 173
Donacia pubicollis Crotch (1873:21) (not Suffrian, 1872:21). Synonymy with Donacia pubicollis Suffrian by Marx (1957). No specimens of Crotch's species were found at MCZ indi- cated as type, probably because it was not recognized that Crotch had also described the species, until Marx (1957). I presume the type must be lost, but it is hardly necessary to designate a neotype for such a well-known species. Donacia (Donaciomima) Medvedev (1973:876). The subgenus Donaciomima was originally based on an eastern Palaearctic species, but was disregarded in subsequent treatments. All Nearctic species of Donacia were formerly placed in Donacia s. str., but Askevold (1990) revalidated Donacia (Donaciomima), and defined it to include the majority of species hitherto placed in the nominate subgenus. Most Nearctic species (23) of Dona- cia are included, of which two remain undescribed. D. assimilis Lacordaire (1845:153). Holotype 0, BMNH. Lacor- daire had before him a single specimen of undetermined sex: "TYPE [red] I TYPE [white] I E. Coll. L(afert6 I Donacia assimilis type Lacord. I Donacia assimilis Dej. h. in Ameri. bor. J. Leconte".
Donacia glabrata Schaeffer (1919:313) (not Solsky, 1872). Based on single 0, USNM holotype: "Type I Clementon, VIII. 6. N.J. I Brooklyn Museum Colln. 1929 I TYPE USNM 42387 I D. glabrata type Schffr. I Donacia assimilis Lac." Synonymy with D. assimilis by Schaeffer (1925). D. biimpressa Melsheimer (1847:159). Lectotype 0, MCZ #33365. Melsheimer had before him at least two specimens; the single remaining specimen is here designated as lectotype: "Melsh. biimpres I Type [red] 1 biimpressa [large Melsheimer label, handwritten] 1 LECTOTYPE 0 Donacia biimpressa Melsheimer design. I.S. Askevold 1990 [red] I MCZ. Lecto- type 33365 [red]".
Donacia aurichalcea Melsheimer (1 847: 159). Evidently based on a single specimen. Holotype 0, MCZ #33364: "Melsh. aurichalcea I Type [red] 1 dissected and remounted IS Askevold 1983 I aurichalcea [large handwritten Melsheimer label] I Donacia biimpressa Melsheimer 1847 det. I.S.



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174 Psyche [Val. 98
Askevold 1983 I M.C.Z. Lectotype 33364 [red]". Synonymy with D. biimpressa by Schaeffer (1925).
Donacia torosa LeConte (1 85 1 :3 13). LeConte did not specify number of specimens, but three specimens found in the LeConte collection are here designated as types. Lectotype (3, MCZ: "pink disc I LECTOTYPE [blue trimmed disc] I TYPE 4243 [red] I D. torosa Lee. I J. LECONTE COLLECTION I LECTOTYPE (3 Donacia torosa LeConte 185 1 designated IS Askevold 1983 [red]". Paralectotype (7: "pink disc I Type 4243 [red] 1 torosa 2 I J. LECONTE COLLECTION 1 PAR- ALECTOTYPE (3 Donacia torosa LeConte [red]"; paralecto- type [sex undetermined, abdomen missing] "pink disc I torosa 3 I J. LECONTE COLLECTION I PARALECTO- TYPE Donacia torosa LeConte [red]". Synonymy with D. biimpressa by Schaeffer (1 925).
D. caerulea Olivier (1795: 10). Neotype (3, CNC #19492, desig- nated by Askevold (1987b:347). Discussed by Askevold (1987b), this species was formerly known as D. aequalis; D. caerulea was used incorrectly by Schaeffer (1925) and Marx (1957), which they applied to certain specimens of D. prox- ima.
Donacia aequalis Say (1 824:248). Neotype (3, MCZ #32948, designated by Askevold (1987b:347); placed in synonymy with D. caerulea by Askevold (1987b).
Donacia confusa Lacordaire (1 845; 109) (not LeConte, 185 1). Lectotype (7, BMNH, designated by Askevold (1987b:347); placed in synonymy with D. caerulea by Askevold (1987b). Donacia confusa LeConte (1 85 1 :3 13) (not Lacordaire, 1845). Lectotype (3, MCZ #32949, designated by Askevold (1987b:347); placed in synonymy with D. caerulea by Askevold (1 987b).
D. cazieri Marx (1957:213). Holotype (3, CNC, allotype 9, CNC, four paratopotypes, CNC, and one paratopotype, AMNH, Goose Bay, Labrador (various dates and collectors, see Marx 1957).
D. confluenta Say (1827:293). Neotype (3, MCZ #32956, desig- nated by Aske~old (1987a:634); removed from synonymy with D. subtilis Kunze by Askevold (1987a).



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D. cuprea Kirby (1 837~225) (not Melsheimer, 1847). Lectotype Q, BMNH, designated, and removed from synonymy with D. quadricollis Say, by Askevold (1987a1633). Donacia curticollis Knab (1905: 122) (not Haupt, 1956). Removed from synonymy of D. quadricollis Say by Askevold (1987a). Lectotype Q, and three paratypes, INHS; designation and details given by Frison (1927: 160). D. dissimilis Schaeffer (1925: 11 5). Holotype Q, USNM: "Florida ex Liebeck 1 Charles Schaeffer Collection 1 HS Barber Bequest 1950 1 dissimilis Holotype C. Schaeffer coll. 1935 Barber 1 Donacia dissimilis Schffr.". Allotype 0, PUR: "Dunedin [Florida] 4.16.24 W.S.B. coll. 1 2724 1 0 Allotype 1 Donacia dissimilis 0 allotype Schaeffer". D. distincta LeConte (1 85 1 :3 13). LeConte did not state the number or sex specimens before him; two specimens were found in the LeConte collection. Lectotype 0 and paralectotype Q, MCZ #4242, here designated: ''0 [pale blue disc] I LECTO- TYPE [blue trimmed disc] 1 Type 4242 [red] 1 D. distincta Lec. aeaualis Kirby 1 LECTOTYPE 0 Donacia distincta LeConte 185 1 designated 1983 IS Askevold [red]" [specimen dissected, with genitalia placed in microvial pinned beneath specimen]; paralectotype Q, "Q [pale blue disc] 1 Type 4242 [red] 1 distincta 2 I J. LECONTE COLLECTION 1 PAR- ALECTOTYPE Q Donacia distincta LeConte [red]". Donacia distincta occidentalis Mead (193811 15). Holotype 0, CAS #4691, allotype Q, CAS #4692, and paratype, MCZ [e.x Fall Coll.], Luthers Pass, Cal. vi-15-1930 AT McClay. Syn- onymy with D. distincta by Marx (19571, to which I agree. D. fulgens LeConte (18511312). Lectotype 0, MCZ #4240, desig- nated by Askevold (1987a:634). At the time of this designa- tion, another syntype specimen was not sent to me; I here add a Q paralectotype from the LeConte collection: "pale blue disc, two cuts 1 Type #4240 [red] I D. fulgens u. 1 J. LECONTE COLLECTION 1 PARALECTOTYPE Q Donacia fulgens LeConte [red]". Removed from synonymy of D. sub- tilis Kunze by Brivio and Balsbaugh (1984). D. hirticollis Kirby (18371226). Holotype 0, BMNH. Kirby stated he had before him a single specimen, which bears the follow- ing labels: "TYPE [red trimmed disc] 1 N. Amer. 5954a



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Psyche [Val. 98
[white disc] 1 Donacia hirticollis Kirby N. Ameri. 5954 Rev. Wm. Kirby [type, on underside]". The specimen lacks the right fore leg and left hind leg, and right elytron smashed by pin.
Donacia rudicollis Lacordaire, 1845: 108. LECTOTYPE Q, BMNH, by present designation. Lacordaire stated he had before him two, supposedly female? specimens. The single specimen found is here designated: "LECTOTYPE [blue trimmed disc] 1 TYPE [red trimmed disc] 1 TYPE [white] 1 E. Coll. Laferti 1 [green, square paper, 4 mm] 1 67.56 1 Donacia rudicollis Type Lac. 1 Donacia rudicollis. Reiche Amer. bor. 1 LECTOTYPE Q Donacia rudicollis Lacordaire 1845 desig- nated lO.vii.1984 I.S. Askevold [red] 1 Donacia hirticollis Kirby det. I.S. Askevold 1984". I have not determined the original source of synonymy.
D. liebecki Schaeffer (1919:314). Holotype 0, USNM #42390: "Type 0 1 Wyandanch L Isld. 1 Brooklyn Museum Colln. 1929 1 liebecki type USNM 42390 [red] 1 Donacia liebecki Schffr. [red bordered]". Allotype 9: "Type Q Allotype I Wyandanch L Isld. 1 liebecki Schffr. Allotype USNM 42390 [red] 1 Brooklyn Museum Colln. 1929". Five paratypes MCZ #27228 (two ex Fall Colln.), Malaga 1.8 N.J. One specimen labelled as a paratype (USNM) may be a paratype, though it does not bear labels indicating it is part of the original syn- type series of 11 specimens: "Lakehurst NJ 1 Ed. A. Bischoff Collection 1 Brooklyn Museum Colln. 1929 I liebecki USNM paratype ? 42390 [red]". Another specimen labelled as a paratype (USNM) cannot be a paratype because it is from Wading River, Long Island, a locality not mentioned by Schaeffer.
Donacia pallipes Lacordaire (1 845: 149) (not Kunze, 18 18). Holotype Q, BMNH. Lacordaire had before him a single specimen which he supposed to be male. It is female? how- ever. "TYPE [red trimmed disc] l TYPE [white] l 67.56 l D. pallipes Lac. 0 type 1 Donacia type 32. pallipes Lac. 119 39 Philadelp Am. Sept. Boston D. Green [green, folded] 1 Dona- cia liebecki Schaeffer det. IS Askevold 1985." D. pallipes was listed in early catalogues as a synonym of D. aequalis Say; Schaeffer (1925) realized that D, pallipes Lac. was a



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19911 Askevold 177
senior synonym of D. liebecki, but preoccupied, thus D. liebecki remains valid. Synonymy with D. liebecki by Schaeffer (1 925).
D. limonia Schaeffer (1925). This name was described as a variety of D. biimpressa Melsh.; Marx (1957) reduced it to syn- onymy with D. biimpressa. I here remove it from synonymy, and recognize D. limonia as a valid species on the basis of distinctive structure of male genitalia; further treatment of this species as a member of the D. distincta-Group is planned. NEW STATUS
D. biimpressa var. limonia Schaeffer (1 925: 100). Holotype 0, USNM: "Indian Pass N.Y. VII. 11. 1 Essex Co. Quirsfeld 1 Brooklyn Museum Coll. 1929 1 limonia Schffr. TYPE USNM 42388 1 Donacia biimpressa limonia Schffr,". Allotype Q, USNM, Van Cortlandt Park, N.Y. (additional details not recorded), and five paratypes, USNM (label details not recorded).
D. magnifica LeConte (1851:310). LeConte did not state the num- ber of specimens before him, but stated he had both male and female specimens. Four specimens in the Leconte collection bear appropriate label data, and are here designated, lecto- type 0, Mcz #4237, one 0 and two QQ paralectotypes, McZ #4237: "LECTOTYPE [blue trimmed disc] 1 0 [pale blue disc] 1 Type 4237 [red] 1 Donacia magnifica h. 1 LECTO- TYPE 0 Donacia magnifica LeConte 1851 designated I.S. Askevold 1984 [red]"; the three paralectotypes are labelled "magnifica 2", "rnagnifica 3" and "magnifica 4", each with the labels "Type 4237" [red] and "PARALECTOTYFE Donacia magnifica LeConte [red]". In earlier catalogues, D. magnifica was listed under D. cineticornis. D. megacornis Blatchley (19 10: 1103). Lectotype 0, PUR. Blatch- ley did not state the number of specimens before him, but designated a lectotype 0 later (Blatchley 1930): "Lake Co., Ind. W.S.B. 5-28-1903 1 Donacia megacornis, sp. nov. [hand- written]".
Donacia megatocera [sic] Weise (1 9 13: 1 18). Unnecessary replacement name. Clavareau (1 9 13123) altered the spelling of megatocera to rnegalocera, probably supposing that it was a printer's error. Weise proposed the name D. megalocera to



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Psyche [Val. 98
replace Blatchley's D. megacornis, on the basis that mega- cornis is a name formed from two languages; the name megacornis is valid, and Schaeffer (1925) restored use of the name D. megacornis, and synonymized D. megatocera. D. porosicollis Lacordaire (1 845: 150). Holotype Q, BMNH. Lacor- daire described this species from a single, apparently female specimen (which it is): "TYPE [red trimmed disc] 1 TYPE [white] 1 Donacia Chvt longipennis parraceicollis Lac. type America bor. Boston D. Greene 627 [green] 1 67.56 1 E. Coll. Chevt. 1 21. [folded paper] 1 D. porosicollis Lac. type". The specimen is a little crushed and is missing the right middle leg.
D. pubescens LeConte (1868:55). Described from a unique speci- men, which LeConte stated was from the Smoky Hill River. Holotype Q, MCZ #4234: "Type [red] 1 D. pubescens Lec.". D. rugosa LeConte (1 878:4l5). Described from a unique speci- men, from Enterprise, Florida. Holotype Q, MCZ #4241: "Enterprise May 20 Fla. 1 402 1 Type 4241 [red] 1 D. rugosa &. 1 J. LECONTE COLLECTION",
D. subtilis Kunze (1818: 12) (replacement name for D. aenea Ahrens).
Donacia aenea Ahrens (1810:21) (not Hoppe 1795:44; not Gis- tel 1857524). Neotype Oy CMP; designation and discussion by Askevold (1987a:633).
Donacia quadricollis Say (1827:282). Neotype 0, MCZ #3295lY designated by Askevold (1987a:633). The name D. quadri- collis was formerly used by authors (e.g. Schaeffer 1925, Wilcox 1954? Marx 19571, applied to the species now known as D. cuprea Kirby (see above) but was synonymized with D. subtilis by Askevold 1987a).
Donacia aerea Lacordaire (1 845: 148) (not Schrank, 1798). Lec- totype 0, BMNH, designated by Askevold (1987a). I have not determined the original source of synonymy. Donacia subtilis magistrigata Mead (1 938: 1 13)- Holotype 0, CAS #4689, allotype Q, CAS #4690, two paratypes? CAS? and one paratype, MCZ #27230 (ex Fall Colln.), Clarksburg? Cal, June-4- 193 1, A.T. McClay. Synonymized with D. ful- gens by Marx (1957) but transferred to synonymy with D. subtilis by Askevold (1987a).




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Donacia tuberculata Lacordaire (1 845: 155). Lectotype 0 and paralectotype Q, BMNH. Two specimens were found that appeared to be authentic Lacordaire specimens, which are here designated: "LECTOTYPE [blue trimmed disc] 1 D. tuberculata Lac 1 67.56 1 LECTOTYPE 0 Donacia tubercu- lata Lacordaire 1845 designated lO.vii.1984 I.S. Askevold [red]". A female is labelled: "PARALECTOTYPE Q [pale blue-trimmed disc] 1 TYPE [red trimmed disc] 1 TYPE [white] 1 E. Coll. Lafert6 1 Donacia tuberculata mihi h. in Ameri. bor. D. Leconte [green, folded] 1 67.56 1 PARALEC- TOTYPE Q Donacia tuberculata Lacordaire 1845 designated lO.vii.1984 I.S. Askevold [red]." Donacia tuberculata was synonymized with D. rufa Say by Schaeffer (19251, an arrangement followed by Marx (1957). Leng (1891) had used D. rufa in the sense used above, as a species of Plateumaris to which it was transferred by Askevold (1991). With trans- ferral of D. rufa to Plateumaris, D. tuberculata becomes the senior, available name for the taxon previously recog- nized as D. rufa, sensu Schaeffer (1925) and Marx (1957). NEW STATUS
Donacia rutila Melsheimer (18471160). Lectotype Q, MCZ #32952, by present designation: "LECTOTYPE [blue trimmed disc] 1 Melsh. rutila 1 M.C.Z. Lectotype 32952 [red] 1 tuberculata [large handwritten Melsh. label] 1 LECTOTYPE Q Donacia rutila Melsheimer, design. I.S. Askevold 1990 [red]". Earlier authors (e.g. Leng 1891) placed D. rutila under D. tuberculata. However, this name was transferred to synonymy of D. rufa. Because D. tuberculata is revalidated, D. rutila becomes its synonym. NEW SYNONYMY D. tuberculifrons Schaeffer (19 19:3 15). Holotype 0, USNM: "Toronto V11.7 Can. 1 Brooklyn Museum Coll. 1929 I TYPE USNM 42389 1 D. tuberculifrons type Schffr.". Allotype Q and one paratype, USNM (label details not recorded). Four paratypes, MCZ #27229 (1 ex Fall Colln.): "Toronto, Can. VII. 1".
This species was treated as a member of the D. subtilis- Group by Askevold (1987a).
D. vicina Lacordaire (1 845: 154) (not Haupt, 1956). Holotype Q, BMNH. Lacordaire described the species from a single



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180 Psyche [vo~. 98
specimen received from Reiche, incorrectly stating it to be male: "HOLOTYPE [red trimmed disc] I TYPE [white] 1 E. Coll. Lafertk I Donacia vicina Dej. h. in Amer. bor. D. Leconte [green]".
Donacia (Donacia) Fabricius, 1775. All Nearctic species of Donacia were formerly placed in Donacia s. str. (e.g. Leng 189 1, Jacoby and Clavareau 1904, Clavareau 19 13, Jolivet 1970, Borowiec 1984), but Askevold (1990) restricted the nominate subgenus to include 10 Nearctic species. D. cincticornis Newman (1838:391). Lectotype 9, BMNH, by pres- ent designation. Newman did not state the number of speci- mens before him. The single specimen remaining is therefore here designated: "LECTOTYPE [blue trimmed disc] I TYPE H.T. [red trimmed disc] I Ent. Club. 44- 12 I R. Foster Tren- ton Falls, New York / Donacia cincticornis type Newm. / LECTOTYPE 9 Donacia cincticornis Newman 1838 desig- nated 1 1 .vii. 1984 I.S. Askevold 1984". Donacia lucida Lacordaire (1 845: 106). Lectotype, BMNH, by present designation. The species was described from at least two specimens. The single remaining specimen under this name is selected as lectotype, which bears the following labels: "LECTOTYPE [blue trim disc] 1 TYPE [red trim disc] 1 Donacia Lucida Lac Type I TYPE 1 E. Coll. Chevt. I D. lucida c? Lac. type / Donacia type 626 lucida (Chvt) Lac. mon 6 Am. bor. Boston D. Greene [green label] I 67.56 I LECTO- TYPE [abdomen missing, probably male] Donacia lucida Lacordaire 1845 designated 10.VII.1984 I.S. Askevold [red] I Donacia cincticornis Newman det. I.S. Askevold 1984". Syn- onymy with D, cincticornis was apparently first by Leng (1 891).
Donacia rufipennis Lacordaire (1 845: 1 10). Lectotype (3, BMNH, by present designation. Lacordaire described this species from four specimens. A male is here selected as lec- totype, bearing the following labels: LECTOTYPE [blue trimmed disc, added by me] I TYPE [red] I Donacia rufipen- nis Type Lac. [white] I TYPE [white] I Donacia type picipen- nis Chevt rufipennis Lac. mon. Ameri. bor. Boston D. Greene [green] / 12 1 67.56 / LECTOTYPE Donacia rufipennis



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Lacordaire 1845 designated 10.VII. 1984 I.S. Askevold [red] / Donacia cincticornis Newman det. I.S. Askevold 1984". Synonymy with D. lucida by Crotch (1873), therefore placed as synonym of D. cincticornis by later authors. Donacia pulchella LeConte (1 85 1 :3 12). LeConte apparently described a single specimen, stating at least that he had no males. The single specimen found in the LeConte collection is therefore assumed to be the holotype, which LeConte incorrectly sexed, Holotype Q, MCZ #4235: "pink disc / Type 4235 [red] 1 D. pulchella LeC.". Synonymy with D. lucida by Crotch (1873), therefore placed as synonym of D. cincticor- nis by later authors.
Donacia antillarum Suffrian (1 864:282). Suffrian described this from at least two specimens, because he gave a range of size. A single specimen here designated as lectotype was found in MLU, associated with the Suffrian label: "Antillarum m. Cuba [large, green, black-bordered label]". LECTOTYPE (sex undetermined, abdomen missing): "25004. [red hand- writing on small label] / LECTOTYPE Donacia antillarum Suffrian 1864 design. I.S. Askevold 1991". Treated as a sub- species of D. cincticornis by Schaeffer (1925); synonymy with D. cincticornis by Marx (1957).
Donacia cincticornis var. tryphera Schaeffer (1925:81). Holo- type 0, USNM: "Centerton VII N.J. / Brooklyn Museum Coll. 1929 / tryphera TYPE USNM 42385". Allotype Q, and 11 paratypes, USNM (label data not recorded). Marx (1957) reduced Schaeffer's subspecies to synonymy; I agree with this synonymy.
Donacia cincticornis var. tenuis Schaeffer (1 925 : 82). Holotype 0, USNM: "Watershield / Swansea SO 11 Aug. 11 Fredk. Knab. / Type c? / tenuis TYPE USNM 42386 / D. cincticornis tenuis Schffr.". Allotype Q, and 16 paratypes, USNM (label data not recorded). Two paratypes, MCZ #27232 (ex Fall Colln.): "Spencer L., N.Y. viii.26.17, H. Dietrich". Marx (1957) reduced Schaeffer's subspecies to synonymy; I agree with this synonymy.
D. edentata Schaeffer (1 9 19:3 13). Holotype a, USNM: "Crum Lk. N.Y. / U.S.N.M. Type No. 42383 / D. edentata type Schffr.". Allotype 9 and five paratypes, USNM (label data not



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182 Psyche [vo~. 98
recorded). One paratype CAS, and eight paratypes, MCZ, "Centerton N.J., 5.3 1".
D. hypoleuca Lacordaire, 1845: 101. Lectotype 0, BMNH, by pres- ent designation. Lacordaire described this species from two, supposedly 9, specimens; however, there is only a single o specimen under hypoleuca which bears an original Lacor- daire label: "Lectotype 0 [blue trim disc added by me] I E. Coll. Laferte I 67.56 I LECTOTYPE c? Donacia hypoleuca Lacordaire 1845 designated 1 1 .VII. 1984 I.S. Askevold [red, added by me]". Also in the BMNH is another specimen (0) which bears a type label; it bears the labels "TYPE [white] 1 This specimen cannot possibly be the type of hypoleuca Lac. The labels have evidently been misplaced. C.G. I Donacia hypoleuca. Van Wintheim h. in Amer. bor. Neu Orleans D. v. Wintheim [green Lacordaire label]". This latter label is evi- dently genuine, according to information given by Lacor- daire, mentioning van Wintheim specifically (1845:lOl); however, the specimen does not agree with the description at all, and is in fact a specimen of D. palmata Olivier. Whether the specimen is a syntype or not is irrelevant, for the other specimen is here designated lectotype in the interest of con- serving the traditional use of D. hypoleuca. D. militaris Lacordaire (1845:115). Lectotype 0, BMNH, by pres- ent designation. Lacordaire did not state the number of speci- mens before him, only that he had no female specimens; a (3 specimen was found: "LECTOTYPE [blue trimmed disc] I TYPE [red trimmed disc] I Donacia militaris type Lac. I TYPE [white] I E. Coll. Chevt. 1 67.56 I D. militaris Lac. [??I type I Donacia type 630 bispinosa (Chvt) 9 militaris Lac mon. America bor. Boston D. Greene [green, folded] I LEC- TOTYPE Donacia militaris Lacordaire 1845 designated 10.vii. 1984 I.S. Askevold [red]". Formerly misidentified as a synonym of D. palmata, but removed from synonymy by Schaeffer (1 925).
Donacia floridae Leng (1 89 1 : 166). No type specimen specifi- cally identified was found in USNM (R.E. White, pers. comm.); however, a specimen, USNM type #558 was found among specimens in "regular collection", which is assumed



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to be a paratype. Synonymized by Schaeffer (1925), which is doubtless correct.
D. palmata Olivier (1795:8). Type(s) almost certainly destroyed; no neotype is designated because Olivier provided a figure of the protarsus of male specimens; the species is unique among Donaciinae in this structure.
Donacia claudicans Germar (1 82 1 : 173). Germar evidently described this from a single male specimen, specifically dis- cussing the unusual and unmistakable protarsal structure. According to Suffrian (1872), he had the Germar collection in his possession (see extended discussion of P. nitida by Askevold, 1991). Thus, Germar's type, if existing, should be in the Suffrian collection. Three specimens (00) are associ- ated with the large, green, black-bordered Suffrian label in MLU: "palmata Oliv. Lec. claudicans Grm. Illinois". Neither of these fits especially the specific colours that Germar described, and so it is quite likely that none of these is the Germar specimen. However, it is not necessary to designate a neotype because the species is so distinctive and Germar described the distinguishing character. I have not traced the source of original synonymization.
Donacia palmata angustipes Marx (1957:237). Holotype 0 and allotype Q #8452, and 19 paratypes, CAS, "Devils Riv. near Del Rio, Texas, vi-16-32, J.O. Martin"; 2 paratypes, CAS, "Devils Riv. near Del Rio, Texas, v-25-32, J.O. Martin". These Texas specimens differ little from other, typical D. palmata from other regions, differing essentially only in that males have a less broadly palmate basal protarsomere. NEW SYNONYMY
D. parvidens Schaeffer (1 9 19:3 1 1). Holotype 0, USNM: "Lahaway Ocean Co. V. N.J. / Brooklyn Museum Coll. 1929 I U.S.N.M. Holotype 42384 / D. parvidens type Schffr.". Four paratypes, USNM (label data not recorded) and one paratype, MCZ, "Lahaway, Ocean Co., N.Y."
D. piscatrix Lacordaire (1 845: 113). Lectotype 0, BMNH, by pres- ent designation. Lacordaire did not state the number of speci- mens before him, only that he had no male specimens. One Q specimen was found: "LECTOTYPE [blue trimmed disc] / TYPE [red] 1 Donacia piscatrix Lacord. 1 E. Coll. Lafertk I D.



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184 Psyche [vo~. 98
piscatrix Lac. / Donacia Am. bor. [green] / 67.56 / LECTO- TYPE 9 Donacia piscatrix Lacordaire 1845 designated 10.vii.1984 I.S. Askevold [red]".
Donacia carolina Lacordaire (1 845: 1 14). Lectotype 0, BMNH. Lacordaire stated he had both a male and female before him. The single, apparently authentic, Lacordaire specimen found among specimens of D. piscatrix, is here designated: "LEC- TOTYPE [blue trimmed disc] / TYPE [red] / Donacia car- olina Lacord. / TYPE [white] / 67.56 / Donacia carolina mihi. h. in Amer. bor. D. Latreille [green] / Donacia piscatrix det. I.S. Askevold '84". I did not trace the original source of syn- onymization.
Donacia cuprea Melsheimer (1 847: 158) (not Kirby, 1837). Lec- totype 0, MCZ, by present designation. Described from at least two specimens, for Melsheimer gave a size range and indicated a "var. a". The single remaining specimen is here designated: "0 [discoloured square] / Melsh. cuprea / Type [red] 1 cupria [sic] [large handwritten Melsheimer label] / LECTOTYPE 0 Donacia cuprea Melsheimer design. I.S. Askevold 1990 [red] / M.C.Z. Lectotype 33366 [red]". Syn- onymy with D. piscatrix apparently first by Schaeffer (1925). Donacia alutacea LeConte (1 85 1 :3 1 1). LeConte had at least two specimens before him. Two specimens found in the LeConte collection are here designated: lectotype 0, MCZ #4239, "LECTOTYPE [blue trimmed disc] / Type [partly red] / o [pink disc] 1 Type 4239 [red] 1 D. alutacea &. cuprea /I Mels. / LECTOTYPE c? Donacia alutacea LeConte 185 1 designated I.S. Askevold 1984 [red]"; and paralectotype 0, "Q [pink disc] / Type 4239 [red] 1 alutacea 2 / PARALECTO- TYPE 0 Donacia alutacea LeConte [red]". Synonymy with D. piscatrix by Crotch (19873).
Donacia congener LeConte (1 85 1 :3 10). LeConte did not state num- ber of specimens before him, only that he had no males. Two specimens in the LeConte collection bear orange discs, so it appears he had at least two specimens, and I here designate them. Lectotype Q, MCZ #4238: "LECTOTYPE [blue trimmed disc] I Q [orange disc] / Type 4238 [red] / D. congener &. 1 LECTOTYPE Q Donacia congener LeConte 185 1 designated I.S. Askevold 1984"; and one 9 paralectotype, MCZ #4238 "9



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19911 Askevold 185
[orange disc] 1 Type 4238 [red] I congener 2 I PARALECTO- TYPE Q Donacia congener LeConte [red]". Synonymy with D. piscatrix by Crotch (1873), treated as subspecies by Schaeffer (1925), resynonymized by Marx (1957).
D. proxima Kirby (1837:225). Lectotype Q, BMNH, designated by Askevold (1987b:347).
Donacia episcopalis Lacordaire (1845: 105). Lectotype 0, BMNH, designated by Askevold (1987b:348). Formerly placed in syn- onymy with D. proxima, transferred to synonymy with D. caerulea by Schaeffer (1925), who used this name used incor- rectly; D. episcopalis was transferred back to junior synonymy with D. proxima by Askevold (1987b).
Donacia californica LeConte (1 861 :357). Holotype Q, MCZ #4236 (cf. Askevold 1987b:348). Given subspecies status by Marx (1957), but this classification rejected by Askevold (1987b). Donacia caerulea, sensu Schaeffer (1925) and Marx (1957). See Askevold (1987b) for discussion.
D. rufescens Lacordaire (1 845: 1 12). Lectotype 0, BMNH, by present designation. Lacordaire had before him two specimens, suppos- edly female. However, both are male, of which one is here des- ignated: "LECTOTYPE [blue trimmed disc] I TYPE [red trimmed disc] I TYPE [white] I Donacia type 629 calcarata Chvt 8 rufescens Lac. mon. America bor. Boston D. Greene [green, folded] I 15.1 67.56 I LECTOTYPE c? Donacia rufescens Lacor- daire 1845 designated 10.vii.1984 I.S. Askevold [red]". D. texana Crotch (1873:22). Lectotype Q, MCZ #5032, by present des- ignation. Crotch did not state the number of specimens before him, but three specimens already indicated as types were found in the MCZ. Lectotype Q, MCZ #5032, "Tex. I Type 5032 [red] 1 D. texana Cr. I LECTOTYPE Q Donacia texana Crotch design. IS Askevold 1990 [red]"; paralectotype Q, MCZ #5032, "Tex. I Type 5032 [red] I texana 2 I PARALECTOTYPE Donacia tex- ana Crotch [red]" and paralectotype 0, MCZ #5032, "Tex. I Type 5032 [red] I texana 3 1 PARALECTOTYPE Donacia tex- ana Crotch [red]".
Donacia texana var. minor Schaeffer (1925:72). Holotype 0, USNM: "Centerton V. 28. N.J. I Donacia texana minor Schffr. (3 I U.S.N.M. Type No. 42382". Allotype Q, and 6 paratypes, USNM (label data not recorded). Marx (1957) did not recognize



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186 Psyche [vo~. 98
D. minor as a subspecies and therefore synonymized D. minor, to which I agree.
Neohaemonia Szekessy (1941:148). This genus was formerly consid- ered congeneric with the Palaearctic genus, Macroplea Samouelle. Revision of genus and establishment of genus status by Askevold (1988).
N. flagellata Askevold (1988:390). Holotype 0, CNC, and 72 paratypes of various locality data deposited in CARR, CAS, CNC, CUIC, EGRC, ISAC, JBWM, ROM, UAE, UMMA, USNM.
N. flohri (Jacoby, 1884: 126). Holotype Q, MCZ #8468. Details given by Askevold (1988).
N. melsheimeri (Lacordaire, 1845:211). Holotype (3, BMNH. Details given by Askevold (1988).
N. minnesotensis Askevold (1988:388). Holotype 0, CNC #19495, and 38 paratypes with various locality data deposited in AMNH, CAS, CMP, CNC, DEFW, EGRC, FMNH, FSCA, ISAC, JBWM, LSUC, MCZ, NMDC, SCSU, UMMA, USNM. N. nigricornis (Kirby, 1837:222). Holotype 9, BMNH. Details given by Askevold (1988).
Haemonia americana Gukrin-Mkneville (1844:259). Holotype 9, BMNH. Details given by Askevold (1988).
I owe the many curators of museums cited above profuse thanks for their inestimable help with my studies about Donaciinae. Without their assistance, this and other publications on this group would suffer sig- nificant incompleteness. Mostly I examined all types myself, but Richard E. White is owed particular thanks for providing details of Schaeffer and Leng types that I was neglectful in recording data about personally, or did not personally examine. Research over a number of years, and costs of publication, were supported by NSERC grant A0428 held by R.E. Roughley, University of Manitoba, and in part also by an NSERC PGS Scholarship, an Ernst Mayr Grant in aid of research at MCZ, and funds provided by CAS to



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19911 Askevold 187
research at that institution. Some costs were supported by a grant from CSRS, USDA to Florida A. & M. University. The Nearctic species of Donaciinae are catalogued according to the most recent classification of its members at tribal, generic and subgeneric levels. Annotations are provided where significant depar- tures have been made from their preceding classification. Donacia tuberculata Lacordaire is removed from synonymy with D. rufa Say (now placed in Plateumaris). Donacia limonia Schaeffer is removed from synonymy with D. biimpressa Melsheimer. Donacia angustipes Marx is reduced to synonymy of D. palmata Olivier; D. rutila Melsheimer is synonymized with D. tuberculata Lacordaire. Details of label data, number, kind and location of preservation of type speci- mens (if existing) of all described taxa are given. Lectotypes (and par- alectotypes, if any) are designated for the following nominal taxa: D. alutacea LeConte, D. biimpressa Melsheimer, D. Carolina Lacordaire, D. cincticornis Newman, D. congener LeConte, D. cuprea Melsheimer, D. distincta LeConte, D. hypoleuca Lacordaire, D. lucida Lacordaire, D. magnifica LeConte, D. militaris Lacordaire, D. pisca- trix Lacordaire, D. rudicollis Lacordaire, D. rufescens Lacordaire, D. rufipennis Lacordaire, D rutila Melsheimer, D. texana Crotch, D. torosa LeConte and D. tuberculata Lacordaire. Types of D. palmata Olivier and D. claudicans Germar are believed lost, but are distinctive taxa whose identity is not in doubt.
AHRENS, A.
1810. Beitrage zu einer Monographie der Rohrkafer. Neue Schriften der natur- forschenden Gesellschaft zu Halle 1: 9-48. ASKEVOLD, I.S.
1987a. The identity of Donacia cuprea Kirby, 1837 and Donacia quadricollis Say, 1827, with a taxonomic revision of members of the Donacia subtilis Kunze - group (Coleoptera: Chrysomelidae: Donaciinae). The Canadian Entomologist 119: 629-645.
1987b. The identity of Donacia caerulea Olivier, 1795 (Coleoptera: Chrysomeli- dae: Donaciinae). The Coleopterists' Bulletin 41: 345-349. 1988. The genus Neohaemonia Szi5kessy in North America (Coleoptera: Chrysomelidae: Donaciinae): systematics, reconstructed phylogeny and geographic history. Transaction of the American Entomological Society 113: 360-430.




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1990. Reconstructed phylogeny and reclassification of the genera of Donaciinae (Coleoptera: Chrysomelidae). Quaestiones Entomologicae 26: 601-664. 1991. Classification, reconstructed phylogeny and geographic history of the New World members of Plateumaris Thomson, 1859 (Coleoptera: Chrysomeli- dae). Memoirs of the Canadian Entomological Society 157: 1-175. BECKER, E.C.
1974. Designation of lectotypes for 13 species of Coleoptera described by Couper in 1864 and 1865. Annales de la Soci6t6 entomologique de Quebec 19: 62-73.
BLATCHLEY, W. S.
1910. An illustrated descriptive catalogue of the Coleoptera or Beetles (exclusive of the Rhynchophora) known to occur in Indiana. Nature Publ. Co., Indi- anapolis. 1386 pp.
1930. Blatchleyana. A list of published writings of W.S. Blatchley, A.B., A.M., L.L.D. of Indianapolis, Indiana and Dunedin, Florida. Together with a chronology of his life: the fixation of types of new genera and species described by him, etc., etc. Nature Publ. Co., Indianapolis. 77 pp. BOROWIEC, L.
1984. Zoogeographical study on Donaciinae of the world (Coleoptera, Chrysomelidae). Polski Pismo Entomologiczne. 53: 433-518. BRIVIO, C. AND E.U. BALSBAUGH, JR.
1984. New genitalic characters for distinguishing Donacia subtilis Kunze and D. fulgens LeConte (Coleoptera: Chrysomelidae). Coleopts. Bull. 38: 305-3 1 1.
CHEN, S.H.
1941. Notes on donaciine beetles. Sinensia 12: 1-17. CLARK, H.
1866. A catalogue of Phytophaga (Coleoptera, Pseudotetramera). Appendix. Williams & Morgate, London and Edinburgh. 1 + 88 pp. CLAVAREAU, H.
1913. Sagrinae, Donaciinae and Criocerinae, In Junk, W. and S. Schenkling, Edi- tors. Coleopterorum Catalogus 51: 1-103. COUPER, W.
1864. Paper V. List of Coleoptera and Diptera taken at Quebec, and other parts of Lower Canada. Transactions of the Library and Historical Society of Que- bec. New Series, Part 2. Session of 1863-1864: 75-93. CROTCH, G.R.
1873. Materials for the study of the Phytophaga of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 25: 19-83. FABRICIUS, J.C.
1775. Systema Entomologicae, sistens insectorum classes, ordines, genera, species, adjectis, synonymis, locis, descriptionibus, observationibus. Korte, Flensburgi et Lipsiae. 8. [30] + 832 pp [not seen]. PRISON, T.H.
1937. A list of the insect types in the collections of the Illinois State Natural His- tory Survey and the University of Illinois. Bulletin of the Illinois Natural History Survey 16: 137-309.
GERMAR, E.F.
181 1. 11. Nachtrage zu Ahrens Monographic der Rohrkafer. Neue Schriften der Naturforschenden Gesellschaft zu Halle I: 26-38.



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19911 Askevold 189
1821. In Wiedemann, C.R.W. and E.F. Germar. Neue exotische Kafer beschrieben von Wiedemann und Germar. Magazin der Entomologie 4: 107-183.
GISTEL, J.N.F.X.
1857. Achthundert und zwanzig neue oder unbeschriebene wirbellose Thiere. Vacuna 2: 513-606. [Also published separately, Straubing, Schorner, 1857. 8: 94 pp, under the title "Vacuna oder die Geheimnisse aus der organischen und leblosen Welt." The former is usually cited.] GOECKE, H.
193 1. Die Gattung Sominella Jocobson (Col. Chrys. Donaciini) (1. Beitrag zur Kenntnis der Donaciinen). Entomologische Blatter fur Biologie und Sys- tematik der Kafer. 27: 152-1 61, 1 plate. GUERIN-MENEVILLE, F.E.
1944. Iconographie du Rkgne Animal de G. Cuvier, representation d'aprks nature de l'une des espkces les plus remarquables et souvent non encore figurees, de chaque genre d'animaux. Tome 111. Paris, J.B. Bailliere, 1829-1844. 8. 576 pp, 104 plates.
HATCH, M.H.
1938. A new species of Donacia from Washington. Pan-Pacific Entomologist 14: 110-112.
1971. The Beetles of the Pacific Northwest Part V. Rhipiceroidea, Sternoxi, Phy- tophaga, Rhynchophora, and Lamellicornia. University of Washington Publications in Biology Vol. 16. xiv + 662 pp. HAUPT, H.
1956. Beitrag zur Kenntnis der eozanen Arthropodenfauna des Geiseltales. Nova Acta Leopoldina (N.F.) 18: 90 pp.
HOPPE, D.H.
1795. Enumeration Insectorum elytratorum circa Erlangam indigenarum secun- dum systema Fabricianum observationibus iconibusque illustrata. Erlan- gae, Palm 1795. iv + 70 pp + 1 pi.
JACOBY, M.
Beschreibung neuer Phytophagen. Stettiner entomologische Zeitung 45: 126-128.
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Coleoptera Phytophaga Fam. Donacidae. In P. Wytsman, Genera Insecto- rum 21: 1-14, 1 plate.
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190 Psyche [vo~. 98
KUNZE, G.
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1845. Monographie des colkoptixes subpentamkres de la famille des phy- tophages. Tome 1. Mimoires de la SociM Royale des Sciences de Liege. Tome 3 (pt. 1). xiii + 740 pp.
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1851. Synopsis of the species of Donacia (Fabr.) inhabiting the United States. Proceedings of the California Academy of Sciences 5: 3 10-3 16. 1857 (1 860). No. 1. Report upon the insects collected on the Survey, pp 1-72 In, Report of Explorations and Surveys, to ascertain the most practicable and economical route for a railroad from the Mississippi River to the Pacific Ocean, made under the direction of the Secretary of War in 1853-5. Vol- ume XI1 Book 11. Washington, T.H. Ford 1860. 1861. New species of Coleoptera inhabiting the Pacific district of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 1861: 338-359.
1868. New Coleoptera collected on the Survey for the extension of the Union Pacific Railway, E.D. from Kansas to Fort Craig, New Mexico. Transac- tions of the American Entomological Society 2: 49-64. 1878. Additional descriptions of new species. Pp. 373-434 In, J.L. LeConte and E.A Schwarz, Editors. Coleoptera of Florida and Michigan. Proceedings of the American Philosophical Society 1878: 353-669. LENG, C.W.
1891. Revision of the Donaciae of Boreal America. Transactions of the American Entomological Society 18: 159-176.
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1843. Beitrag zur Kaefer-Fauna der Aleutischen Inseln, der Insel Sitkha und Neu- Californiens. Bulletin de la Sociitk Impiriale des Naturalistes de Moscou 16: 175-314.
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192 Psyche [vo~. 98
INDEX TO SPECIES, GENUS AND FAMILY GROUP NAMES aenea, 178
aerea, 178
aequalis, 174, 176
afinis, 17 1
alutacea, 184
americana, 186
angustipes, 183
antillarum, 18 1
assimilis, 173
aurichalcea, 173
aurifera, 168
balli, 168
biimpressa, 173, 177
binodosa, 170
caerulea, 174, 185
californica, 185
carolina, 184
cataractae, 169, 170
cazieri, 174
chalcea, 17 1
cincticornis, 170,
177, 180, 181
claudicans, 183
confluenta, 174
confusa, 174
congener, 184
cuprea, 175, 1 84
curticollis, 175
dissimilis, 175
distincta, I75
diversa, 168
dives, 169
Donacia (Donacia),
180
Donacia (Dona-
ciomima), 173
Donaciella, 172
Donaciini, 172
dubia, 168, 169
edentata, 18 1
emarginata, 169, 170
episcopalis, 1 85
femoralis, 169
flagellata, 186
flavipennis, 169
flavipes, 169, 171
flohri, 186
floridae, 182
frosti, 169
fulgens, 175, 178
fulvipes, 169
gentilis, 169
germari, 169
glabrata, 173
harrisii, 172
hirticollis, 175
hypoleuca, 182
idola, 169
indica, 169
juncina, 170
jucunda, 17 1
kirbyi, 171
liebecki, 176
limonia, 177
lodingi, 171
longicollis, 170
lucida, 180, 181
Macroplea, 186
magistrigata, 178
magnified, 177
megacornis, 177
megalocera, 177
megatocera, 177
melsheimeri, 186
metallica, 169
militaris, 1 82
minnesotensis, 186
minor, 185, 186
nana, 169
Neohaemonia, 1 86
neomexicana, 170
nigricornis, 186
nitida, 170, 171
notmani, 170
occidentalis, 175
vacifica. 170
pallipes, 176
palmata, 182, 183
parva, 169
parvidens, 183
piscatrix, 183, 184
Plateumarini, 168
Plateumaris, 168
Poecilocera, 172
porosicollis, 178
proxima, 174, 185
pubescens, 178
pubicollis, 172
pulchella, 1 8 1
pusilla, 170, 171
pyritosa, 17 1
quadricollis, 175,
178
robusta, 171
rudicollis, 176
rufa, 170, 171, 179
rufescens, 185
rufipennis, 180
rugifrons, 170
rugosa, 178
rutila, 179
serricauda, 169
schaeferi, 170, 171
shoemakeri, 169, 171
Sominella, 172
subtilis, 174, 175,
178
sulcicollis, 17 1
tenuis, 181
texana, 1 85
torosa, 174
tryphera, 18 1
tuberculata, 17 1, 179
tuberculifrons, 179
vermiculata, 170
vicina, 179
wallisi, 169




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