Studies on North American Carboniferous insects. 9. A new species of Eubleptidae from Mazon Creek (Palaeodictyoptera).
Psyche 99(2-3):147-152, 1992.
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STUDIES ON NORTH AMERICAN CARBONIFEROUS
INSECTS. 9. A NEW SPECIES OF EUBLEPTIDAE FROM MAZON CREEK (PALAEODICTYOPTERA)
Museum of Comparative Zoology
Harvard University, Cambridge, Mass. 02138 In a previous part of this series (Carpenter, 1983) I discussed the structure and relationships of the extinct family Eubleptidae and its unique generic representative, Eubleptus danielsi Hand- lirsch. This was based on my study of the 17 specimens of the insect then known, all from the Upper Carboniferous of Mazon Creek, Illinois. Since then, Mr. Michael A. Madsen, of Darien, Illi- nois, has kindly loaned me for study a well-preserved specimen of a new species also belonging to Eubleptus. Since this contributes considerably to our understanding of the family, I am describing it here.
Family Eubleptidae Handlirsch, 1906
Eubleptus maculosus n. sp.
This species is similar to dmielsi, so far as known. Both species have the same basic pattern of venation in fore and hind wings, but in maculosus the wings are more slender and the points of origin of RS and of the forking of M are much nearer the center line of the wings. Also, MA in both fore and hind wings is unbranched in maculosus but deeply forked in danielsi. Of special interest is the presence of four large, dark, submarginal spots extending to mid- wing in both fore and hind wings of maculosus, in addition to cer- tain other spots along other main veins. In danielsi both wings have four dark spots on the anterior margins, but none on the rest of the wings.
Very little of the body is preserved. The abdomen is 11.5 mm. long and 1.5 mm. wide; the cerci, which are very prominent in Manuscript received 30 July 1992.
148 Psyche [VOI. 99
Figure 1. Eublcpt~~ marufosus, n. sp. Dorsal view of type specimen, obverse half. Wing expanse, 32 mm,
danielsi, are not clearly preserved in maculosus, and the head is not preserved at all. The fore and hind wings are 14 rnm. long. Holotype: in the collection of Michael A. Madsen, Darien, Illi- nois. The fossil was collected by him at Mazon Creek, Pit 11, Braidwood, Illinois (Upper Carboniferous), and includes both counterparts.
19921 Carpenter-Eubleptidae 149
Figure 2. Eubleptus maculosus, n.sp. (type), showing dark spots. The family Eubleptidae was proposed by Handlirsch in 1906 for a single, fragmented specimen of a new species, Eubleptus danielsi, from the Upper Carboniferous of Mazon Creek, Illinois. As noted by Handlirsch, this was the smallest Carboniferous insect known, and it still stands as one of the smallest pterygote insects found in Paleozoic deposits. Handlirsch placed the family in the order Palaeodictyoptera, although the fragmentary nature of the fossil gave little evidence of its phylogenetic position. In 191 1 Handlirsch added another new genus and species (Athymodictya parva from Mazon Creek) to the family, but that has subsequently been shown to be a synonym of danielsi (Carpenter, 3 983). The systematic position of the family Eubleptidae has been doubtful for many years. Martynov mentioned his uncertainty about its ordinal position in 3 938, and Laurentiaux in 1953 placed the family in a new order, Eubleptidia, but without giving a defini- tion of the order. Rohdendorf accepted that order in 1962, although he changed the name to Eubleptodia. A few years later (19651, fol- lowing my examination of the two previously mentioned speci- mens, I concluded that the insects were most like the members of the order Palaeodictyoptera as Handtirsch thought. In 1983 I was
150 Psyche [Vol. 99
Figure 4. Eubleptus maculosus, n. sp. (type). Venational diagram of fore and hind wings.
able to study 17 recently collected specimens of danielsi from Mazon Creek, most of which had been found by private collectors. Many of the fossils were very well preserved. From my study of them I concluded that the new evidence supported Handlirsch's original assumption that they belonged to the order Palaeodicty- optera, and were, in fact, close relatives of the family Spilapteridae (Carpenter, 1983). The structure of the new species, maculosus, supports that conclusion.
Studies on North American Carboniferous insects. 4. The genera Metropator, Eubleptus, Hapaloptera, and Hadentomum. (Palaeodicty- optera). Psyche, 72: 175-190.
Studies on North American Carboniferous insects. 7. The structure and relationships of Eubleptus danielsi (Palaeodictyoptera). Psyche, 90: 81-95.
l906a. Revision of American Palaeozoic insects. Proceedings of the U.S. National Museum, 29: 661-820.
1906b.Die fossilen Insekten und die Phylogenie der rezenten Formen. Ein Handbuch fur Palaontologen un Zoologen. Engelmann, Leipsig. p. 1 -640.
1911. New Paleozoic insects from the vicinity of Mazon Creek, Illinois. American Journal of Science (4) 21: 297-326, 353-377. LAURENTIAUX, D.
1953. Classe des insectes. In Trait6 de Paleontologic (ed., Piveteau), p. 397-527.
MARTYNOV, A. V.
1938. Etudes sur l'histoire geologique et de phylogenie des ordres des insects. Trudy paleontologicheskogo instituta academii nauk SSSR, 7: 1-150.
ROHDENDORF, B. B.
1962. Osnovy Palaeontologii: Tracheata, Mandibulata. Akad. Nauk., USSR, p. 1-374.
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